Information on EC 3.2.1.73 - licheninase

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The expected taxonomic range for this enzyme is: Eukaryota, Bacteria

EC NUMBER
COMMENTARY hide
3.2.1.73
-
RECOMMENDED NAME
GeneOntology No.
licheninase
REACTION
REACTION DIAGRAM
COMMENTARY hide
ORGANISM
UNIPROT
LITERATURE
Hydrolysis of (1->4)-beta-D-glucosidic linkages in beta-D-glucans containing (1->3)- and (1->4)-bonds
show the reaction diagram
REACTION TYPE
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
hydrolysis of O-glycosyl bond
-
-
-
-
SYSTEMATIC NAME
IUBMB Comments
(1->3)-(1->4)-beta-D-glucan 4-glucanohydrolase
Acts on lichenin and cereal beta-D-glucans, but not on beta-D-glucans containing only 1,3- or 1,4-bonds.
CAS REGISTRY NUMBER
COMMENTARY hide
37288-51-0
-
ORGANISM
COMMENTARY hide
LITERATURE
UNIPROT
SEQUENCE DB
SOURCE
strain A-25
-
-
Manually annotated by BRENDA team
strain EGW039 (CGMCC 0635)
-
-
Manually annotated by BRENDA team
strain EGW039(CGMCC 0635)
-
-
Manually annotated by BRENDA team
isozymes EG1 and EG2
-
-
Manually annotated by BRENDA team
Bacillus sp. KCCM 90078
-
-
-
Manually annotated by BRENDA team
gene bg1314
UniProt
Manually annotated by BRENDA team
gene bglS
UniProt
Manually annotated by BRENDA team
strain A3
-
-
Manually annotated by BRENDA team
strain GN156
-
-
Manually annotated by BRENDA team
gene SU40-glu
UniProt
Manually annotated by BRENDA team
isolated from Asian giant hornet, Vespa mandarinia, nest, gene bgl5-1
UniProt
Manually annotated by BRENDA team
isolated from Asian giant hornet, Vespa mandarinia, nest, gene bgl5-1
UniProt
Manually annotated by BRENDA team
-
-
-
Manually annotated by BRENDA team
Bispora sp.
-
UniProt
Manually annotated by BRENDA team
var. miniatus
-
-
Manually annotated by BRENDA team
i.e. Malbranchea sulfurea
UniProt
Manually annotated by BRENDA team
i.e. Malbranchea sulfurea
UniProt
Manually annotated by BRENDA team
Orpinomyces sp.
strain PC-2
-
-
Manually annotated by BRENDA team
OsEGL1; cv. Yukihikari, gene OsEGL1
UniProt
Manually annotated by BRENDA team
strain GS01
-
-
Manually annotated by BRENDA team
gene plicA
UniProt
Manually annotated by BRENDA team
a hypercellulolytic mutant strain
-
-
Manually annotated by BRENDA team
a hypercellulolytic mutant strain
-
-
Manually annotated by BRENDA team
variant microsporus
-
-
Manually annotated by BRENDA team
strain ZJL4, gene LicB
-
-
Manually annotated by BRENDA team
cultivar Musketeer
Uniprot
Manually annotated by BRENDA team
Thermomonospora sp.
-
-
-
Manually annotated by BRENDA team
strain MSB8
-
-
Manually annotated by BRENDA team
strain ZJU-T
-
-
Manually annotated by BRENDA team
strain ZJU-T
-
-
Manually annotated by BRENDA team
gene Zobellia_2431
UniProt
Manually annotated by BRENDA team
gene Zobellia_2431
UniProt
Manually annotated by BRENDA team
GENERAL INFORMATION
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
evolution
physiological function
additional information
SUBSTRATE
PRODUCT                       
REACTION DIAGRAM
ORGANISM
UNIPROT
COMMENTARY
(Substrate) hide
LITERATURE
(Substrate)
COMMENTARY
(Product) hide
LITERATURE
(Product)
Reversibility
r=reversible
ir=irreversible
?=not specified
(1-3,1-4)-beta-D-glucan + H2O
?
show the reaction diagram
-
i.e. barley (1-3,1-4)-beta-glucan
-
-
?
1,3-1,4-beta-D-glucan + H2O
?
show the reaction diagram
-
-
-
-
?
1,3-1,4-beta-glucan + H2O
?
show the reaction diagram
-
-
-
-
?
2,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
2,4-dinitrophenol + beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
3,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
3,4-dinitrophenol + beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl 3-O-beta-cellobiosyl-beta-D-glucopyranoside + H2O
4-methylumbelliferone + 3-O-beta-cellobiosyl-beta-D-glucopyranoside
show the reaction diagram
-
-
-
-
?
4-methylumbelliferyl 3-O-beta-cellobiosyl-beta-D-glucoside
4-methylumbelliferone + 3-O-beta-cellobiosyl-beta-D-glucose
show the reaction diagram
-
-
-
-
?
4-methylumbelliferyl beta-D-cellobioside + beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl beta-D-cellobioside + beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl beta-D-cellobioside + beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl cellobiose + H2O
?
show the reaction diagram
-
-
-
-
?
4-methylumbelliferyl laminaribiose + H2O
?
show the reaction diagram
-
-
-
-
?
4-methylumbelliferyl-(1,3)-beta-D-glucooligosaccharides + H2O
4-methylumbelliferol + beta-D-oligosaccharides
show the reaction diagram
-
-
-
-
?
4-methylumbelliferyl-beta-D-Gal-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
?
show the reaction diagram
-
Gal substrate has a 1.3fold higher kcat/KM-value than Glc substrate
-
-
r
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc + H2O
4-methylumbelliferone + beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
4-methylumbelliferone + beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
?
show the reaction diagram
-
Gal substrate has a 1.3fold higher kcat/KM-value than Glc substrate
-
-
r
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
4-methylumbelliferone + beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + H2O
4-methylumbelliferone + beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
show the reaction diagram
-
-
-
-
-
4-nitrophenyl beta-D-cellobioside + H2O
?
show the reaction diagram
-
1% of the activity with carboxymethyl-cellulose
-
-
?
4-nitrophenyl beta-D-glucopyranosyl-(1-4)-beta-D-glucopyranosyl-(1-4)-beta-D-glucopyranosyl-(1-3)-2-deoxy-beta-D-erythro-pentopyranoside + H2O
?
show the reaction diagram
-
2-deoxy analog is better substrate than corresponding 2-hydroxy substrate
-
-
?
4-nitrophenyl beta-D-glucopyranosyl-(1-4)-beta-D-glucopyranosyl-(1-4)-beta-D-glucopyranosyl-(1-3)-beta-D-arabinopyranoside + H2O
?
show the reaction diagram
-
2-deoxy analog is better substrate than corresponding 2-hydroxy substrate
-
-
?
alpha-laminaribiosyl fluoride + 4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-methylumbelliferyl-beta-D-glucopyranoside
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-methylumbelliferyl-beta-D-xylopyranoside
4-methylumbelliferyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Xyl + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-nitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc
4-nitrophenyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-nitrophenyl-beta-D-glucopyranoside
4-nitrophenyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + 4-nitrophenyl-beta-D-xylopyranoside
4-nitrophenyl-beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Xyl + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + methyl beta-D-Glc-(1-3)-beta-D-Glc
methyl beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
alpha-laminaribiosyl fluoride + methyl beta-D-Glc-(1-4)-beta-D-Glc
methyl beta-D-Glc-(1-3)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc + F-
show the reaction diagram
-
-
-
-
-
arabinoxylan + H2O
?
show the reaction diagram
-
-
-
-
?
avicel + H2O
?
show the reaction diagram
AZO blue beta-glucan + H2O
?
show the reaction diagram
-
-
-
-
?
barley beta-D-glucan + H2O
?
show the reaction diagram
barley beta-glucan + H2O
3-O-beta-cellobiosyl-D-glucose + 3-O-beta-cellotriosyl-D-glucose
show the reaction diagram
barley beta-glucan + H2O
3-O-beta-cellobiosyl-D-glucose + 3-O-beta-cellotriosyl-D-glucose + ?
show the reaction diagram
-
-
-
-
?
barley beta-glucan + H2O
?
show the reaction diagram
barley beta-glucan + H2O
beta-glucan oligosaccharides
show the reaction diagram
barley beta-glucan + H2O
cellotriose + cellotetraose + cellobiose + D-glucose
show the reaction diagram
barley beta-glucan + H2O
Glc-beta(1,4)-Glc-beta(1,3)-Glc-beta(1,4)-Glc + Glc-beta(1,3)-Glc-beta(1,4)-Glc-OH
show the reaction diagram
barley-beta-glucan + H2O
?
show the reaction diagram
beta-(1-3),(1-4)-D-glucan + H2O
beta-D-glucose + ?
show the reaction diagram
-
-
-
-
?
beta-1,3-1,4-glucan + H2O
3-O-cellobiosyl-D-glucose + 3-O-cellotriosyl-D-glucose
show the reaction diagram
beta-1,3-1,4-glucan + H2O
?
show the reaction diagram
beta-1,3/1,4-glucan + H2O
?
show the reaction diagram
beta-D-glucan + H2O
?
show the reaction diagram
beta-glucan + H2O
?
show the reaction diagram
-
-
-
-
?
carboxymethyl cellulose + H2O
?
show the reaction diagram
carboxymethyl-cellulose + H2O
?
show the reaction diagram
carboxymethylcellulose + H2O
?
show the reaction diagram
cellohexaose + H2O
cellobiose + cellotriose
show the reaction diagram
-
no release of D-glucose
-
?
cellopentaose + H2O
cellobiose + D-glucose
show the reaction diagram
cellulose + H2O
?
show the reaction diagram
chitosan + H2O
?
show the reaction diagram
81.3% of the activity with barley beta-D-glucan
-
-
?
CM-cellulose + H2O
?
show the reaction diagram
KM079629
low activity
-
-
?
Glc-beta(1,4)-Glc-beta(1,4)-Glc-beta(1,3)-Glc + H2O
cellobiose + laminaribiose
show the reaction diagram
Glcbeta(1-3)Glcbeta(1-4)Glcbeta(1-4)Glcbeta(1-4)Glc + H2O
glucose + Glcbeta(1-3)Glcbeta(1-4)Glcbeta(1-4)Glc
show the reaction diagram
-
-
-
-
?
Glcbeta(1-4)Glcbeta(1-4)Glcbeta(1-3)Glcbeta(1-4)Glcbeta(1-4)Glc-OMe + H2O
?
show the reaction diagram
-
-
-
-
?
Glcbeta(1-4)Glcbeta(1-4)Glcbeta(1-4)Glcbeta(1-3)Glc + H2O
?
show the reaction diagram
-
-
-
-
?
Glcbeta(1-4)Glcbeta(1-4)Glcbeta(1-4)Glcbeta-O-4-methylumbeliferone + H2O
?
show the reaction diagram
-
-
-
-
?
Glcbeta3Glcbeta-methylumbelliferone + H2O
?
show the reaction diagram
-
-
-
-
?
glucan tetrasaccharide + H2O
D-glucose + glucan trisaccharide
show the reaction diagram
containing two beta-1,4-linkages separated by one beta-1,3-linkage, i.e. G4G3G4G
containing one beta-1,4-linkage separated by one beta-1,3-linkage, i.e. G4G3G, which is not further degraded
-
?
hydroxyethylcellulose + H2O
?
show the reaction diagram
-
-
-
?
laminarin + H2O
?
show the reaction diagram
laminarin + H2O
D-glucose + ?
show the reaction diagram
-
-
-
?
laminaritriose + H2O
D-glucose + laminaribiose
show the reaction diagram
the smallest oligosaccharide that can be degraded by the enzyme
-
-
?
lichenan + H2O
3-O-beta-cellobiosyl-D-glucose + 3-O-beta-cellotriosyl-D-glucose
show the reaction diagram
lichenan + H2O
?
show the reaction diagram
lichenan + H2O
cellobiose + Glc-beta(1,4)-Glc-beta(1,3)-Glc-beta(1,4)-Glc + Glc-beta(1,3)-Glc-beta(1,4)-Glc-OH
show the reaction diagram
lichenan + H2O
cellotriose
show the reaction diagram
lichenan + H2O
cellotriose + cellotetraose
show the reaction diagram
lichenan + H2O
D-glucose + ?
show the reaction diagram
lichenan + H2O
lichenan oligosaccharides
show the reaction diagram
-
-
-
-
?
lichenan + H2O
reducing sugar + ?
show the reaction diagram
lichenin + H2O
?
show the reaction diagram
oat beta-glucan + H2O
?
show the reaction diagram
soluble starch + H2O
?
show the reaction diagram
KM079629
low activity
-
-
?
starch + H2O
?
show the reaction diagram
wheat arabinoxylan + H2O
?
show the reaction diagram
xylan + H2O
?
show the reaction diagram
xylan + H2O
xylose + ?
show the reaction diagram
Thermomonospora sp.
-
-
-
-
?
barley beta-glucan + H2O
additional information
-
NATURAL SUBSTRATES
NATURAL PRODUCTS
REACTION DIAGRAM
ORGANISM
UNIPROT
COMMENTARY
(Substrate) hide
LITERATURE
(Substrate)
COMMENTARY
(Product) hide
LITERATURE
(Product)
REVERSIBILITY
r=reversible
ir=irreversible
?=not specified
barley beta-glucan + H2O
?
show the reaction diagram
barley-beta-glucan + H2O
?
show the reaction diagram
KM079629
-
-
-
?
beta-1,3-1,4-glucan + H2O
3-O-cellobiosyl-D-glucose + 3-O-cellotriosyl-D-glucose
show the reaction diagram
beta-1,3-1,4-glucan + H2O
?
show the reaction diagram
laminarin + H2O
?
show the reaction diagram
G0L5X4
-
-
-
?
lichenan + H2O
?
show the reaction diagram
oat beta-glucan + H2O
?
show the reaction diagram
additional information
-
-
METALS and IONS
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
Fe3+
-
89.6% residual activity at 12 mM
Li+
-
103.25% activity at 1 mM
NaCl
1.5fold activation at 0.5 M, the enzyme is acid-, alkali- and salt-tolerant, the purified recombinant His-tagged enzyme retains over 90% activity in 4 M NaCl at 25°C for 24 h
Triton X-100
about 20% activation
additional information
INHIBITORS
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
IMAGE
(3,4)-epoxybutyl beta-D-cellobioside
-
identification of Glu105 at the active site
1-ethyl-3-(3-dimethylaminopropyl)carbodiimide
-
-
2,4,6-Trinitrobenzenesulfonic acid
Thermomonospora sp.
-
-
2,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
-
2-mercaptoethanol
3,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
-
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
-
5,5'-dithiobis 2-nitrobenzoic acid
Thermomonospora sp.
-
-
Ba2+
-
8% inhibition at 1 mM
beta-mercaptoethanol
-
inhibits the enzyme activity at 4 mM by 82.7%
CaCl2
-
10 mM, 33% inhibition
compatibility of EG1 and EG2 lichenases with solid laundry detergents
-
-
-
dithiothreitol
-
1 mM, 21% inhibition
epoxyalkyl cellobiosides
-
Glc-beta-1,3-isofagomine
-
-
glucono-delta-lactone
-
-
H2O2
-
at 0.5% w/v hydrogen peroxide, isozymes EG1 and EG2 retain 82.8% and 90.28% of their activities, respectively. At 1% w/v hydrogen peroxide, isozyme EG2 retains 60% activity, while isozyme EG1 is completely inhibited
imidazole
-
competitive
K+
-
95.61% residual activity at 1 mM
Na+
-
inhibits the enzyme activity at 4 mM by 47.3%; inhibits the enzyme activity at 4 mM by 80.3%
NaCl
-
30%, halflife of wild-type 230 h, mutant K48A 694 h, mutant K48L 693 h, respectively
Ni2+
-
inhibits the enzyme activity at 4 mM by 11.5%
o-phthaldehyde
Thermomonospora sp.
-
-
PCMB
-
-
Sodium dodecyl sulfate
Bispora sp.
strong inhibition
sodium perborate
-
isozymes EG1 and EG2 retain 58% and 66.8% of their activities after 1 h of incubation at 40°C in the presence of 0.2% w/v sodium perborate, respectively
-
Sr2+
-
inhibits the enzyme activity at 4 mM by 49%
Tris
-
competitive
Urea
-
14.5% residual activity at 7 mM
additional information
-
ACTIVATING COMPOUND
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
IMAGE
2-mercaptoethanol
slightly activating
Avicel
-
mutants show higher levels of activity compared to wild type
-
beet pulp
-
mutants show higher levels of activity compared to wild type
-
beta-mercaptoethanol
-
0.1%
N-bromosuccinimide
-
-
solk floc
-
mutants show higher levels of activity compared to wild type
-
Triton X-100
activates to 221.30%
Tween-20
activates to 159.20%
Tween-80
activates to 267.73%
wheat bran
-
mutants show higher levels of activity compared to wild type
-
additional information
-
KM VALUE [mM]
SUBSTRATE
ORGANISM
UNIPROT
COMMENTARY hide
LITERATURE
IMAGE
0.7
2,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
pH 7.2, 30°C
0.7
3,4-dinitrophenyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
pH 7.2, 30°C
0.37 - 0.6
4-methylumbelliferyl 3-O-beta-cellobiosyl-beta-D-glucoside
0.058
4-methylumbelliferyl cellobiose
-
-
0.19
4-methylumbelliferyl laminaribiose
-
-
1
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
pH 7.2, 30°C
0.5
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
pH 7.2, 30°C
0.6
4-methylumbelliferyl-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-beta-D-Glc
-
pH 7.2, 30°C
1.36
beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
-
pH 7.0, 35°C
0.34
beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
-
pH 7.0, 35°C
0.15
beta-D-Glc-(1-4)-beta-D-Glc-(1-4)-beta-D-Glc-(1-3)-alpha-D-Glc-fluoride
-
pH 7.0, 35°C
5.3
Glcbeta3Glcbeta-methylumbelliferone
-
pH 7.0, 37°C, 50 mM sodium phosphate buffer
2.29 - 14.7
Laminarin
0.13 - 10.3
lichenan
additional information
Barley beta-glucan