BRENDA - Enzyme Database
show all sequences of 3.4.22.3

Ficin

Liener, I.E.; Friedenson, B.; Methods Enzymol. 19, 261-273 (1970)
No PubMed abstract available

Data extracted from this reference:

Activating Compound
Activating Compound
Commentary
Organism
Structure
EDTA
enhances activity
Ficus sp.
Inhibitors
Inhibitors
Commentary
Organism
Structure
Chloroacetamide
-
Ficus sp.
HgCl2
-
Ficus carica
HgCl2
-
Ficus glabrata
HgCl2
-
Ficus sp.
iodoacetamide
-
Ficus sp.
iodoacetic acid
-
Ficus carica
iodoacetic acid
-
Ficus sp.
NEM
-
Ficus sp.
Molecular Weight [Da]
Molecular Weight [Da]
Molecular Weight Maximum [Da]
Commentary
Organism
26500
-
ultracentrifugation
Ficus anthelmintica
Organism
Organism
UniProt
Commentary
Textmining
Ficus anthelmintica
-
-
-
Ficus carica
-
var. Kadota
-
Ficus glabrata
-
-
-
Ficus sp.
-
-
-
Substrates and Products (Substrate)
Substrates
Commentary Substrates
Literature (Substrates)
Organism
Products
Commentary (Products)
Literature (Products)
Organism (Products)
Reversibility
Substrate Product ID
benzoyl-L-argininamide + H2O
-
95732
Ficus sp.
benzoyl-L-Arg + NH3
-
-
-
?
Collagen + H2O
-
95732
Ficus sp.
?
-
-
-
?
Fibrin + H2O
-
95732
Ficus sp.
?
-
-
-
?
Gelatin + H2O
-
95732
Ficus sp.
?
-
-
-
?
Hemoglobin + H2O
-
95732
Ficus sp.
?
-
-
-
?
hippuryl amide + H2O
-
95732
Ficus sp.
hippuric acid + NH3
-
-
-
?
hippuryl methyl ester + H2O
-
95732
Ficus sp.
hippuric acid + methanol
-
-
-
?
Nalpha-Benzoyl-L-Arg ethyl ester + H2O
-
95732
Ficus sp.
Nalpha-Benzoyl-L-Arg + ethanol
-
-
-
?
Nalpha-Benzoyl-L-Arg ethyl ester + H2O
-
95732
Ficus glabrata
Nalpha-Benzoyl-L-Arg + ethanol
-
-
-
?
p-nitrophenyl-carbobenzoxyglycinate + H2O
-
95732
Ficus sp.
?
-
-
-
?
soy proteins + H2O
-
95732
Ficus sp.
?
-
-
-
?
pH Optimum
pH Optimum Minimum
pH Optimum Maximum
Commentary
Organism
6.5
-
-
Ficus sp.
Activating Compound (protein specific)
Activating Compound
Commentary
Organism
Structure
EDTA
enhances activity
Ficus sp.
Inhibitors (protein specific)
Inhibitors
Commentary
Organism
Structure
Chloroacetamide
-
Ficus sp.
HgCl2
-
Ficus sp.
HgCl2
-
Ficus glabrata
HgCl2
-
Ficus carica
iodoacetamide
-
Ficus sp.
iodoacetic acid
-
Ficus sp.
iodoacetic acid
-
Ficus carica
NEM
-
Ficus sp.
Molecular Weight [Da] (protein specific)
Molecular Weight [Da]
Molecular Weight Maximum [Da]
Commentary
Organism
26500
-
ultracentrifugation
Ficus anthelmintica
Substrates and Products (Substrate) (protein specific)
Substrates
Commentary Substrates
Literature (Substrates)
Organism
Products
Commentary (Products)
Literature (Products)
Organism (Products)
Reversibility
ID
benzoyl-L-argininamide + H2O
-
95732
Ficus sp.
benzoyl-L-Arg + NH3
-
-
-
?
Collagen + H2O
-
95732
Ficus sp.
?
-
-
-
?
Fibrin + H2O
-
95732
Ficus sp.
?
-
-
-
?
Gelatin + H2O
-
95732
Ficus sp.
?
-
-
-
?
Hemoglobin + H2O
-
95732
Ficus sp.
?
-
-
-
?
hippuryl amide + H2O
-
95732
Ficus sp.
hippuric acid + NH3
-
-
-
?
hippuryl methyl ester + H2O
-
95732
Ficus sp.
hippuric acid + methanol
-
-
-
?
Nalpha-Benzoyl-L-Arg ethyl ester + H2O
-
95732
Ficus sp.
Nalpha-Benzoyl-L-Arg + ethanol
-
-
-
?
Nalpha-Benzoyl-L-Arg ethyl ester + H2O
-
95732
Ficus glabrata
Nalpha-Benzoyl-L-Arg + ethanol
-
-
-
?
p-nitrophenyl-carbobenzoxyglycinate + H2O
-
95732
Ficus sp.
?
-
-
-
?
soy proteins + H2O
-
95732
Ficus sp.
?
-
-
-
?
pH Optimum (protein specific)
pH Optimum Minimum
pH Optimum Maximum
Commentary
Organism
6.5
-
-
Ficus sp.
Other publictions for EC 3.4.22.3
No.
1st author
Pub Med
title
organims
journal
volume
pages
year
Activating Compound
Application
Cloned(Commentary)
Crystallization (Commentary)
Engineering
General Stability
Inhibitors
KM Value [mM]
Localization
Metals/Ions
Molecular Weight [Da]
Natural Substrates/ Products (Substrates)
Organic Solvent Stability
Organism
Oxidation Stability
Posttranslational Modification
Purification (Commentary)
Reaction
Renatured (Commentary)
Source Tissue
Specific Activity [micromol/min/mg]
Storage Stability
Substrates and Products (Substrate)
Subunits
Synonyms
Temperature Optimum [°C]
Temperature Range [°C]
Temperature Stability [°C]
Turnover Number [1/s]
pH Optimum
pH Range
pH Stability
Cofactor
Ki Value [mM]
pI Value
IC50 Value
Activating Compound (protein specific)
Application (protein specific)
Cloned(Commentary) (protein specific)
Cofactor (protein specific)
Crystallization (Commentary) (protein specific)
Engineering (protein specific)
General Stability (protein specific)
IC50 Value (protein specific)
Inhibitors (protein specific)
Ki Value [mM] (protein specific)
KM Value [mM] (protein specific)
Localization (protein specific)
Metals/Ions (protein specific)
Molecular Weight [Da] (protein specific)
Natural Substrates/ Products (Substrates) (protein specific)
Organic Solvent Stability (protein specific)
Oxidation Stability (protein specific)
Posttranslational Modification (protein specific)
Purification (Commentary) (protein specific)
Renatured (Commentary) (protein specific)
Source Tissue (protein specific)
Specific Activity [micromol/min/mg] (protein specific)
Storage Stability (protein specific)
Substrates and Products (Substrate) (protein specific)
Subunits (protein specific)
Temperature Optimum [°C] (protein specific)
Temperature Range [°C] (protein specific)
Temperature Stability [°C] (protein specific)
Turnover Number [1/s] (protein specific)
pH Optimum (protein specific)
pH Range (protein specific)
pH Stability (protein specific)
pI Value (protein specific)
Expression
General Information
General Information (protein specific)
Expression (protein specific)
KCat/KM [mM/s]
KCat/KM [mM/s] (protein specific)
752461
Milosevic
Comparative stability of fici ...
Ficus carica
Amino Acids
51
829-838
2019
-
1
-
-
-
-
-
-
-
-
-
-
1
1
-
-
-
-
-
1
-
1
2
-
-
-
-
7
-
1
-
1
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
1
-
1
2
-
-
-
7
-
1
-
1
-
-
-
-
-
-
-
752630
Homaei
-
Purification, catalytic, kine ...
Ficus johannis
Biocatal. Agricult. Biotechnol.
10
360-366
2017
-
-
-
-
-
-
-
1
-
8
1
-
-
1
-
-
1
-
-
1
-
-
1
1
-
1
1
-
1
1
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
8
1
-
-
-
-
1
-
1
-
-
1
1
1
1
-
1
1
1
-
-
-
-
-
-
1
1
755474
Baidamshina
Targeting microbial biofilms ...
Ficus
Sci. Rep.
7
46068
2017
-
1
-
-
-
-
-
-
-
-
-
-
-
13
-
-
-
-
-
-
-
-
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
753868
Zare
Autolysis control and structu ...
Ficus
Int. J. Biol. Macromol.
84
464-471
2016
-
-
-
-
-
1
-
-
-
-
-
-
-
4
-
-
1
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
733039
Haesaerts
Crystallization and preliminar ...
Ficus carica
Acta Crystallogr. Sect. F
71
459-465
2015
-
-
-
1
-
-
-
-
-
-
4
-
-
4
-
-
1
-
-
1
-
-
1
-
4
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
4
-
-
-
-
1
-
1
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
734568
Raskovic
Characterisation of general pr ...
Ficus carica
J. Sci. Food Agric.
96
576-582
2015
-
-
-
-
-
-
-
-
-
-
-
-
-
5
-
-
-
-
-
1
1
-
1
-
1
1
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
1
-
1
-
1
-
-
-
1
-
-
-
-
-
-
-
-
-
755021
Baeyens-Volant
A novel form of ficin from Fi ...
Ficus carica
Phytochemistry
117
154-167
2015
-
-
-
-
-
-
8
-
-
1
2
-
-
4
-
-
1
-
-
3
-
-
5
1
1
1
1
-
-
1
1
-
-
-
-
-
-
-
-
-
-
-
-
-
8
-
-
-
1
2
-
-
-
-
1
-
3
-
-
5
1
1
1
-
-
1
1
-
-
-
-
-
-
-
-
735275
Gagaoua
-
Three-phase partitioning as an ...
Ficus carica
Sep. Purif. Technol.
132
461-467
2014
-
-
-
-
-
-
-
-
-
-
1
-
-
1
-
-
1
-
-
-
-
-
1
-
1
1
-
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
1
-
-
-
-
1
-
1
-
1
-
1
-
-
-
-
-
-
-
-
-
734500
Fu
-
Ficin-catalyzed asymmetric ald ...
Ficus carica
J. Mol. Catal. B
97
1-4
2013
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
16
-
1
2
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
16
-
2
-
-
-
-
-
-
-
-
-
-
-
-
-
734867
Zare
Purification and autolysis of ...
Ficus carica
Phytochemistry
87
16-22
2013
-
-
-
-
-
-
2
-
-
-
-
-
-
6
-
-
1
-
-
2
-
2
1
-
3
1
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
2
-
-
-
-
-
-
-
-
-
1
-
2
-
2
1
-
1
-
-
-
1
-
-
-
-
-
-
-
-
-
718202
Azarkan
Selective and reversible thiol ...
Ficus carica
Phytochemistry
72
1718-1731
2011
-
-
-
-
-
-
6
-
-
-
-
-
-
6
-
-
1
-
-
5
10
-
2
1
5
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
6
-
-
-
-
-
-
-
-
-
1
-
5
10
-
2
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
718344
Devaraj
-
Comparison of activity and con ...
Ficus carica
Process Biochem.
46
458-464
2011
-
-
-
-
-
-
-
-
-
-
-
-
2
1
-
-
-
-
-
1
-
-
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
2
-
-
-
-
1
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
710296
Valdes-Rodriguez
Recombinant amaranth cystatin ...
Ficus sp.
Plant Physiol. Biochem.
48
469-475
2010
-
-
-
-
-
-
1
-
-
-
-
-
-
9
-
-
1
-
-
1
-
-
1
-
1
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
1
1
-
-
-
-
-
-
-
-
1
-
1
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
710654
Sidek
-
Denatured states of ficin indu ...
Ficus sp.
Turk. J. Biochem.
35
45-49
2010
-
-
-
-
-
1
-
-
-
-
-
-
-
1
-
-
1
-
-
1
-
-
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
1
-
1
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
708754
Devaraj
Characterization of acid-induc ...
Ficus carica
Int. J. Biol. Macromol.
45
248-254
2009
-
-
-
-
-
-
-
-
-
-
1
-
-
4
-
-
1
-
-
2
-
-
-
1
1
-
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
1
-
2
-
-
-
1
-
-
-
-
-
-
1
-
-
-
-
-
-
-
677510
Sekizaki
Application of several types o ...
Ficus sp.
Amino Acids
34
149-153
2008
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
1
-
-
10
-
1
-
-
-
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
10
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
698401
Devaraj
Purification, characterization ...
Ficus carica
J. Agric. Food Chem.
56
11417-11423
2008
-
1
-
-
-
1
3
-
-
-
1
1
-
12
-
1
1
-
-
3
1
-
2
1
1
-
-
12
-
1
1
-
-
-
-
-
-
1
-
-
-
-
1
-
3
-
-
-
-
1
1
-
-
1
1
-
3
1
-
2
1
-
-
12
-
1
1
-
-
-
-
-
-
-
-
665093
Ruemenapf
-
434. Cathepsin T ...
Ficus glabrata
Handbook of proteolytic enzymes (Barrett, A. J. , Rawlings, N. D. , Woessner, J. F. , eds. ) Academic Press
2nd Ed.
1404
2004
1
-
-
-
-
-
1
2
1
-
-
1
-
1
-
-
1
1
-
5
-
-
7
1
-
-
-
-
-
1
1
-
-
-
-
-
1
-
-
-
-
-
-
-
1
-
2
1
-
-
1
-
-
-
1
-
5
-
-
7
1
-
-
-
-
1
1
-
-
-
-
-
-
-
-
95731
Hänsler
The Application of papain, fic ...
Ficus carica
J. Pept. Sci.
1
283-287
1995
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
-
-
-
1
-
-
9
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
-
9
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95734
Mazda
Use of standarized protease en ...
Ficus sp.
Transfus. Med.
5
43-50
1995
-
1
-
-
-
-
-
-
-
-
-
-
-
3
-
-
-
-
-
-
-
-
-
-
-
-
-
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-
-
-
-
-
-
-
-
1
-
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-
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-
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-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95733
Patel
Variation in the P2-S2 stereoc ...
Ficus sp.
Biochem. J.
281
553-559
1992
-
-
-
-
-
-
-
-
-
-
-
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-
1
-
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-
-
-
-
-
2
-
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-
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-
-
2
-
-
-
-
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-
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-
-
-
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-
-
95727
Monter
Kinetically controlled synthes ...
Ficus carica
Biotechnol. Appl. Biochem.
14
183-181
1991
-
1
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-
-
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1
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1
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1
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8
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1
1
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1
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1
-
1
-
-
8
-
-
-
-
-
1
1
-
-
-
-
-
-
-
-
95735
Mariani
A new enzymatic method to obta ...
Ficus sp.
Mol. Immunol.
28
69-77
1991
-
1
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-
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4
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-
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-
-
95723
Björk
Interaction between chicken cy ...
Ficus sp.
Biochemistry
29
1770-1776
1990
-
-
-
-
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1
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-
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8
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1
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1
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-
-
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-
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-
-
-
-
-
-
-
-
-
-
-
95710
Lynn
-
Ficin E, a serine-centred prot ...
Ficus elastica
Phytochemistry
25
1559-1561
1986
-
-
-
-
-
-
2
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1
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1
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1
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1
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1
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1
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2
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1
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1
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1
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-
-
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-
1
-
-
-
-
-
-
-
-
-
95711
Minakata
Acidic cysteine proteinase inh ...
Ficus sp.
Biol. Chem. Hoppe-Seyler
366
15-18
1985
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-
-
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1
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3
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1
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1
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1
-
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-
-
-
-
-
-
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-
-
-
-
-
-
30289
Gray
Benzyloxycarbonylphenylalanylc ...
Ficus sp.
Biochem. J.
219
325-328
1984
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-
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3
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11
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11
-
-
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-
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-
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-
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-
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-
-
-
-
81558
Rodis
Naturally occurring protein cr ...
Ficus sp.
Plant Physiol.
74
907-911
1984
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-
-
-
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1
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-
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4
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1
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1
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1
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-
-
-
-
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-
-
-
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-
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-
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-
36125
Wakamatsu
Comparison of properties of th ...
Ficus sp.
J. Biol. Chem.
257
14653-14656
1982
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1
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1
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-
1
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-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95715
Friedenson
Evidence that ficin is a glyco ...
Ficus sp.
Biochim. Biophys. Acta
342
209-212
1974
-
-
-
-
-
-
-
-
-
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1
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2
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2
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-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95716
Anderson
Purification of ficin by affin ...
Ficus sp.
Anal. Biochem.
60
417-423
1974
-
-
-
-
-
-
-
-
-
-
-
-
-
1
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-
1
-
-
1
1
-
-
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-
-
-
-
-
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-
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-
1
-
1
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95717
Hall
Proflavine interactions with p ...
Ficus sp.
Biochemistry
13
2087-2092
1974
-
-
-
-
-
-
2
3
-
-
-
-
-
1
-
-
-
-
-
1
-
-
3
-
-
-
-
-
3
-
-
-
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-
-
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-
2
-
3
-
-
-
-
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-
1
-
-
3
-
-
-
-
3
-
-
-
-
-
-
-
-
-
-
95718
Hall
Proflavine interactions with p ...
Ficus sp.
Biochemistry
13
2082-2087
1974
-
-
-
-
-
-
1
-
-
-
-
-
-
1
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-
1
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-
1
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-
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-
1
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1
-
-
1
-
-
-
-
-
-
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-
-
-
-
-
-
-
-
95719
Kortt
On the specificity and pH depe ...
Ficus sp.
Biochemistry
13
2029-2037
1974
-
1
-
-
-
-
-
24
-
-
-
-
-
1
-
-
-
-
-
-
-
-
17
-
-
-
-
-
24
-
-
-
-
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-
1
-
-
-
-
-
-
-
-
24
-
-
-
-
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-
-
-
-
-
-
17
-
-
-
-
24
-
-
-
-
-
-
-
-
-
-
95720
Kortt
Ficins (EC 3.4.22.3). Purifica ...
Ficus glabrata
Biochemistry
13
2023-2028
1974
-
-
-
1
-
1
1
-
-
-
-
-
-
5
-
-
1
-
-
3
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
1
-
1
-
1
-
-
-
-
-
-
-
-
-
1
-
3
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95721
Sugiura
Studies on proteinases from Fi ...
Ficus carica
Biochim. Biophys. Acta
350
38-47
1974
2
-
-
1
-
-
6
-
-
-
1
-
-
1
-
1
1
-
-
1
-
1
1
1
1
1
-
2
-
1
1
-
-
-
-
-
2
-
-
-
1
-
-
-
6
-
-
-
-
1
-
-
-
1
1
-
1
-
1
1
1
1
-
2
-
1
1
-
-
-
-
-
-
-
-
95722
Sasaki
Interaction of human serum pro ...
Ficus sp.
J. Biochem.
75
171-177
1974
-
-
-
-
-
-
1
-
-
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1
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-
-
-
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1
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-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95725
Whitaker
Ficin- and papain-catalyzed re ...
Ficus glabrata
Arch. Biochem. Biophys.
148
208-216
1972
-
-
-
-
-
-
-
1
-
-
-
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-
1
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-
-
-
-
-
-
2
-
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-
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-
1
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-
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-
-
-
-
2
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95724
Hollaway
The pH-dependence of rates of ...
Ficus sp.
Eur. J. Biochem.
24
332-341
1971
-
-
-
-
-
-
-
5
-
-
-
-
-
1
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-
-
-
1
-
-
1
-
-
-
-
-
4
-
-
-
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-
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-
-
-
-
-
-
-
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-
-
-
5
-
-
-
-
-
-
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-
-
1
-
-
1
-
-
-
-
4
-
-
-
-
-
-
-
-
-
-
95726
Jones
Comparative studies on four su ...
Ficus glabrata
J. Biol. Chem.
245
2765-2772
1970
-
-
-
-
-
-
-
1
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4
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1
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1
1
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-
1
-
-
2
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-
-
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-
-
-
-
-
-
-
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-
-
-
-
-
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-
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-
1
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4
-
-
-
1
1
-
1
-
-
2
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95732
Liener
-
Ficin ...
Ficus anthelmintica, Ficus carica, Ficus glabrata, Ficus sp.
Methods Enzymol.
19
261-273
1970
1
-
-
-
-
-
8
-
-
-
1
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-
4
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-
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11
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-
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-
1
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-
-
-
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-
1
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-
-
-
-
-
-
8
-
-
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1
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-
-
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-
-
-
-
-
11
-
-
-
-
-
1
-
-
-
-
-
-
-
-
-
95728
Williams
Multiple molecular forms of Fi ...
Ficus glabrata
Plant Physiol.
44
1574-1583
1969
-
-
-
-
-
-
-
-
-
-
1
-
-
4
-
-
1
-
-
1
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
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-
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-
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-
1
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-
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1
-
1
1
-
1
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95729
Englund
Studies on ficin. I. Its isola ...
Ficus glabrata
Biochemistry
7
163-175
1968
-
-
-
-
-
-
-
-
-
-
1
-
-
1
-
-
-
-
-
1
-
1
4
-
-
-
-
-
-
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-
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-
-
-
-
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-
1
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-
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1
-
1
4
-
-
-
-
-
-
-
-
-
-
-
-
-
-
-
95730
Metrione
Purification, partial characte ...
Ficus sp.
Arch. Biochem. Biophys.
122
137-143
1967
-
-
-
-
-
-
-
-
-
-
-
-
-
1
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-
1
-
-
-
1
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-
-
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-
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1
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1
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