5.4.99.53 gene MRN1, recombinant expression in Rhodobacter capsulatus strain SB1003. and Synechocystis sp. PCC 6803 using promoters Pnif and PcoaT, coexpression with CAS1, LUP1, and THAS1. Subcloning in Saccharomyces cerevisiae strain GIL77, both marnerol and the putative hydroxymarnerol are detected in the reference yeast strain expressing MRN1. Isolation of triterpenes, while cycloartenol and lupeol represent typical plant triterpenes with tetracyclic plant sterol and pentacyclic plant triterpene scaffolds, respectively, thalianol and marneral exhibit more unusual tri- and monocyclic structures 749117