5.4.99.53 malfunction an Arabidopsis mrn1 knock-out mutant shows round-shaped leaves and late flowering is isolated under long-day conditions. The late-flowering phenotype is much more severe under short-day conditions than under longday conditions. The mrn1 mutant displays aberrant seed morphology, a low germination rate, and root growth retardation. Mrn1 mutant shows a delayed embryo development 730614 5.4.99.53 physiological function the OSC MRN1 from Arabidopsis thaliana catalyzes the formation of the seco-triterpene marneral via an unusual chair-boat substrate conformation, resulting in a further atypical, monocyclic triterpenoid structure, LC-MS analysis 749117