1.1.1.227 evolution the enzyme is a member of the SDR superfamily of enzymes 1.1.1.227 metabolism borneol is converted into camphor by BDH in borneol-degrading strain, Pseudomonas sp. strain TCU-HL1 and is further decomposed through a camphor degradation pathway 1.1.1.227 additional information molecular modeling and docking of the three-dimensional structure of Pseudomonas sp. TCU-HL1 BDH 1.1.1.227 physiological function neither wild-tpe nor a gene deletion mutant is able to grow on (-)-borneol or (-)-camphor as sole carbon source. Both strains grow similarly on (+)-borneol or (+)-camphor