2.4.1.276 2 UDP-glucose + zeaxanthin = 2 UDP + zeaxanthin bis(beta-D-glucoside) - - 2.4.1.276 2 UDP-glucose + zeaxanthin = 2 UDP + zeaxanthin bis(beta-D-glucoside) binding of zeaxanthin is the rate-limiting step for the reaction sequence. The monoglucoside, as an intermediate, never reaches very high levels compared with zeaxanthin 716795