1.2.99.6 carboxylate reductase no activity in Clostridium aceticum 3.5.4.9 methenyltetrahydrofolate cyclohydrolase Clostridium aceticum 4.2.1.39 gluconate dehydratase Clostridium aceticum 6.3.4.3 formate-tetrahydrofolate ligase Clostridium aceticum