1.13.12.8 Watasenia luciferin + O2 bioluminescence 1.13.12.8 Watasenia luciferin + O2 - 1.13.12.8 Watasenia luciferin + O2 coelenterazine disulfate is the Watasenia scintillans luciferin substrate, and the reaction requires ATP, Mg2+ and molecular oxygen. Protein microcrystals in the arm photophores catalyse the bioluminescent reaction using ATP and the substrate coelenterazine disulfate