Reaction catalyzed by (KDO)3-lipid IVA (2-4) 3-deoxy-D-manno-octulosonic acid transferase (2.4.99.15)

		+			=			+
	alpha-Kdo-(2->8)-alpha-Kdo-(2->4)-alpha-Kdo-(2->6)-lipid IVA	+		CMP-beta-Kdo	=		alpha-Kdo-(2->8)-[alpha-Kdo-(2->4)]-alpha-Kdo-(2->4)-alpha-Kdo-(2->6)-lipid IVA	+		CMP