4.2.3.32 levopimaradiene synthase elimination of diphosphate Ginkgo biloba Q947C4 4.2.3.35 syn-pimara-7,15-diene synthase elimination of diphosphate Oryza sativa - 4.2.3.35 syn-pimara-7,15-diene synthase elimination of diphosphate Picea abies Q675L5