1.2.1.26 2,5-dioxovalerate dehydrogenase trans-4-hydroxy-L-proline degradation II 3.5.4.22 1-pyrroline-4-hydroxy-2-carboxylate deaminase trans-4-hydroxy-L-proline degradation II 5.1.1.8 4-hydroxyproline epimerase trans-4-hydroxy-L-proline degradation II