2.4.1.91 flavonol 3-O-glucosyltransferase quercetin triglucoside biosynthesis 2.4.1.239 flavonol-3-O-glucoside glucosyltransferase quercetin triglucoside biosynthesis 2.4.1.240 flavonol-3-O-glycoside glucosyltransferase quercetin triglucoside biosynthesis