1.1.1.219 dihydroflavonol 4-reductase leucopelargonidin and leucocyanidin biosynthesis 1.14.11.9 flavanone 3-dioxygenase leucopelargonidin and leucocyanidin biosynthesis 1.14.14.82 flavonoid 3'-monooxygenase leucopelargonidin and leucocyanidin biosynthesis