2.7.7.24 glucose-1-phosphate thymidylyltransferase dTDP-4-O-demethyl-beta-L-noviose biosynthesis 4.2.1.46 dTDP-glucose 4,6-dehydratase dTDP-4-O-demethyl-beta-L-noviose biosynthesis 5.1.3.13 dTDP-4-dehydrorhamnose 3,5-epimerase dTDP-4-O-demethyl-beta-L-noviose biosynthesis