1.3.1.28 2,3-dihydro-2,3-dihydroxybenzoate dehydrogenase 2,3-dihydroxybenzoate biosynthesis 3.3.2.1 isochorismatase 2,3-dihydroxybenzoate biosynthesis 5.4.4.2 isochorismate synthase 2,3-dihydroxybenzoate biosynthesis