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This enzyme has been characterized from the bacterium Pseudomonas putida KT2440 and is involved in the degradation pathway of syringate and gallate. It catalyses the interconversion of two of the tautomers of 4-oxalomesaconate, a reaction that can also occur spontaneously.
disruption of the galD gene does not prevent growth of the Pseudomonas putida KTGALdgalD mutant strain in 5 mM gallic acid, but the mutant strain grows poorly at gallic acid concentrations lower than 2 mM