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This enzyme has been characterized from the bacterium Pseudomonas putida KT2440 and is involved in the degradation pathway of syringate and 3,4,5-trihydroxybenzoate. It catalyses the interconversion of two of the tautomers of 4-oxalomesaconate, a reaction that can also occur spontaneously.
disruption of the galD gene does not prevent growth of the Pseudomonas putida KTGALdgalD mutant strain in 5 mM gallic acid, but the mutant strain grows poorly at gallic acid concentrations lower than 2 mM