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This enzyme along with EC 184.108.40.206, tropine dehydrogenase, represents a branch point in tropane alkaloid metabolism . Tropine (the product of EC 220.127.116.11) is incorporated into hyoscyamine and scopolamine whereas pseudotropine (the product of EC 18.104.22.168) is the first specific metabolite on the pathway to the calystegines . Both enzymes are always found together in any given tropane-alkaloid-producing species, have a common substrate, tropinone, and are strictly stereospecific .