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This enzyme along with EC 126.96.36.199, tropine dehydrogenase, represents a branch point in tropane alkaloid metabolism . Tropine (the product of EC 188.8.131.52) is incorporated into hyoscyamine and scopolamine whereas pseudotropine (the product of EC 184.108.40.206) is the first specific metabolite on the pathway to the calystegines . Both enzymes are always found together in any given tropane-alkaloid-producing species, have a common substrate, tropinone, and are strictly stereospecific .