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Also oxidizes other tropan-3alpha-ols, but not the corresponding beta-derivatives . This enzyme along with EC 220.127.116.11, tropinone reductase II, represents a branch point in tropane alkaloid metabolism . Tropine (the product of EC 18.104.22.168) is incorporated into hyoscyamine and scopolamine whereas pseudotropine (the product of EC 22.214.171.124) is the first specific metabolite on the pathway to the calystegines . Both enzymes are always found together in any given tropane-alkaloid-producing species, have a common substrate, tropinone, and are strictly stereospecific .